As discussed in the Expanded Basic Pane, some decoders allow operation on Analog or Conventional DC layouts. At the discretion of the manufacturer, some of the non-motor responses may be changeable by setting CV's in the decoder. This pane is where you would set these values. Typical would be whether any of the various functions (such as lights and bell) would be on or off when in DC mode. Consult your decoders manual for options available
Some are very complex and others are very simple, such as alternate analog or DCC only.